PhD program in ecology, Integrated Science Lab (IceLab) & Department of Ecology & Environmental Sciences, Umeå University, Umeå, Sweden (2009-2014)
PhD program in Ecology, Department of Biology, University of New Mexico, Albuquerque, New Mexico, USA (2004-2014) (transferred to Umeå University)
M. S. in Applied Mathematics, Department of Mathematics & Statistics, University of New Mexico, New Mexico, USA (2007-2008)
M. S. in Forest Landscape Ecology, Department of Forest Science, Oregon State University, Corvallis, Oregon, USA (2000-2003)
B. S. in Forest Science & Biological Aspects of Conservation, Department of Forest Ecology & Management, University of Wisconsin-Madison, Wisconsin, USA (1995-2000)
B. A. in English, Kansai University of Foreign Studies, Hirakata, Osaka, Japan (1990-1994)